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*spoilerish*

I always thought that the heads were Ellisime, the queen (I think) of the elven City of Suldanesslar. Jon Irenicus was in love with her, and she is the woman that the clones are made of waaaaay back in the first dungeon in SoA.

If you're playing SoA, you can go to Watcher's Keep any time during chapters 2,3 and 6 (I recommend 6, because some of the battles there are insane). When you actually complete SoA, you will _then_ start the expansion.